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Which of the following can transfer the genes in bacterial nucleoid as well?

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Which of the following can transfer the genes in bacterial nucleoid as well?

(a) F- plasmid

(b) R factor

(c) F’ plasmid

(d) HFR

I have been asked this question in an online quiz.

My query is from Gene Mapping in Bacteria by Conjugation in division Genomics : The Mapping and Sequencing of Genomes of Cytogenetics

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The correct option is (d) HFR

For explanation I would say: While F’ can transfer one or two of the bacterial gene as well, but only HFR can transfer the gene from the nucleoid of the bacteria as it is the F plasmid incorporated in the central chromosome.

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